What is the exact (historical) connection between second-order arithmetic and Hilbert-Bernays' Grundlagen der Mathematik?
Some background: the literature on Reverse Mathematics contains a number of claims regarding the connection between second-order arithmetic and Hilbert-Bernays' Grundlagen der Mathematik. Most of these are vague (some might say inaccurate), probably in part due to the fact that the Grundlagen have not been translated to English (in full).
Here is the gap I wish to see filled:
On one hand, Hilbert-Bernays define a logical system H (for Hilbert) in Grundlagen der Mathematik (Supplement IV) and formalise mathematics therein. Other systems are sketched, as well as the associated development, but no real details are given. System H uses the language of second-order arithmetic enriched with third-order parameters, like symbols for Feferman's $\mu^2$ and a version of Kleene's $\exists^3$.
On the other hand, second-order arithmetic $Z_2$ only has first and second-order variables. Someone must have thrown out the third-order parameters for some reason.
This leads to more specific questions:
Who first introduced $Z_2$ and where (and why)? Who first claimed $Z_2$ can be found in the Grundlagen?