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Why do you expect the normalized operator, as you have defined, to be nonzero? Shouldn't it have a zero at the special representations, i.e. $\chi_1\chi_2^{-1}=p^{\pm1}$? Most probably to get a non-vanishing holomorphic operator one normalizes by $\epsilon(0,\chi_1\chi_2^{-1})\frac{L(1,\chi_2\chi_1^{-1})}{L(0,\chi_1\chi_2^{-1})}.$
Can one say that the Plancherel constant (density) defined by Shahidi is actually analytic continuation of the Plancherel density for the tempered representation?
I might be wrong but I am confused with your "impression". Take $G=SL_2(\mathbb{R})$. The discrete series representations basically consist of various (non-zero) weight modular forms, where as the principal series representations contain Maass forms which are part of the discrete spectrum of $L^2(\Gamma\backslash G)$. So "discrete series representation" is somewhat smaller than "discrete spectrum in $L^2$".
Or in other words how do you estimate sum of Fourier coefficients for groups where you don't have nice Hecke relation like $\rho(n)=\rho(1)\lambda(n)$?