Examples of naturally occurring Quadratic forms or quadrics. I am always fascinated when a quadratic form (or a quadric) arises naturally. I have
some elementary examples, but most of all, I want to learn more examples. I hope this question isn't considered too vague for MO.  Most forms I list are really
elementary, and all are finite dimensional.
I got most of the following examples from M.Berger, Geometry I & II, and from the truly beautiful book "Eléments de géométrie : actions de groupes" by french author Rached Meinmné.
$(0)$ the discriminant on the affine space of unitary degre 2 polynomials
$(i)$ the determinant on endomorphisms of a 2 dimensional vector space, and
$\mathrm{Tr}^2-4\mathrm{det}$
$(ii)$ the radical on the space of quadratic forms on a 2 dimensional vector space,
and the isotrope cone (not sure about the name, degenerate cone?). 
$(iii)$ the family of hermitian forms (built from the Wronskian) on the solution
space of the discrete Schroedinger equation that allow one to show the existence of
right and left side $L^2$ solutions, and the Weyl m function.
$(iv)$ If $\Delta$ is any $2$ dimensional complex vector space, then
$\mathrm{Herm}(\Delta)$, the real vector space of hermitian forms on $\Delta$,
carries a natural quadratic form obtained by constructing an essentially unique
morphism $\rho$ from $\mathrm{Herm}(\Delta)$ to
$\mathrm{Hom}(\Delta\oplus\overline{\Delta})$ such that for all
$h\in\mathrm{Herm}(\Delta),~\rho(h)^2$ is proportional to $\mathrm{Id}$, the proportionality defining the quadratic form. Here, $\rho$ only depends on a choice of
a nonzero element $\omega\in\Lambda^2\Delta^*$.
$(v)$ If $V$ is a 4 dimensional vector space, then $\Lambda^2 V$ carries the natural
quadric $Q(v)=v\wedge v$ where $\Lambda^4 V$ is identified with the underlying
field, which vanishes exactly when $v$ comes from the canonical map
$\mathrm{Gr}(2,V)\rightarrow P\Lambda^2V$.
I remember reading about one on the space of circles, but I forgot the details. What other examples of natural quadratic forms are there? 
 A: Dear Olivier, in line with the more advanced nature of this site, let me give an example of a less elementary nature.
Consider a compact Riemann surface $X$ of genus 2 and on it stable vector bundles $E$ of rank 2 whose determinant bundle $\Lambda ^2E$ is isomorphic to some fixed line bundle $L$ of degree $-1$. Newstead has proved that the moduli space of those vector bundles is the intersection of two quadrics in five-dimesional projective space $\mathbb P^5(\mathbb C)$. And one of those quadrics is the Klein quadric in $\mathbb P^5(\mathbb C)$ parametrizing the lines in some three-dimensional projective space canonically associated to $X$ and $L$. 
 (A Klein quadric is the quadric you mention in number (v) of your list.)
References
P E. Newstead  Stable bundles of rank 2 and odd degree over a curve of genus 2, Topology 7 (1968), 205-215.
For a  geometric description including the role of the Klein quadric, see:
M. S. Narasimhan and S. Ramanan  Moduli of Vector Bundles on a Compact Riemann Surface, 
 Annals of Mathematics, Vol. 89, No. 1, 1969 , pp. 14-51.
A: Two advanced examples from the theory of abelian varieties (for example, elliptic curves) where it is non-trivial to prove that you even get a quadratic form.
Let $A/k$ be an abelian variety defined over an algebraically closed field. Then the degree map
$$ \deg : \operatorname{End}(A)\longrightarrow\mathbb Z $$
is a quadratic form. A reference for this is Mumford's Abelian Varieties (or my Arithmetic of Elliptic Curves for the dimension 1 case).  To indicate why this is non-trivial, I will mention that if you know it for $k=\overline{\mathbb F}_p$, then you can use basic facts about the Frobenius map (roughly, that $a+b\phi$ is separable if $p\nmid a$) to prove the Weil estimate for $\#A(\mathbb F_{p^n})$.
The second example is when $k$ is the algebraic closure of $\mathbb Q$. Let $D$ be a symmetric divisor on $A$. Then the canonical height function
$$ \hat h_D : A(k) \longrightarrow \mathbb R $$
 of Neron and Tate defined by
$$ \hat h_D(P) = \lim_{n\to\infty} 4^{-n}h_D(2^nP) $$
is a quadratic form. Further, if $D$ is ample, then $\hat h_D(P)=0$ if and only $P$ is a point of finite order. Canonical heights are of great importance in studying the arithmetic of abelian varieties. For example, they appear prominently in the statement of the Birch-Swinnerton-Dyer conjecture. They are discussed in Lang's Fundamentals of Diophantine Geometry and my book with Hindry Diophantine Geometry: An Introduction. (Again, the elliptic curve case is covered in my AEC.)
A: In topology, Poincaré duality implies that given a connected and oriented compact manifold M of dimension 4k, the cup product gives rise to an integral non-degenerate symmetric bilinear form on the "middle"  cohomology group $H^{2k}(M,\mathbf Z)$.
This gives rise to the definition of the signature, which will maybe be of interest to you. A variant for 4k+2 dimensional manifolds gives rise to the famous Kervaire invariant. 
See 
 https://en.wikipedia.org/wiki/Signature_%28topology%29
for example.
A Good reference is the book by Milnor and Husemoller. 
A: Quadratic forms occur naturally, as stress energy functions, in Bob Connelly's work on tensegrity structures.
You can find a wealth of details on his website, in particular the book http://www.math.cornell.edu/~web7510/framework.pdf
A: Binary quadratic forms arise in nature as norm forms for a quadratic field. This point of view has various consequences in number theory.


*

*For a fixed negative discriminant (the definite case), Gauss discovered that the quadratic forms (or their $\mathrm{SL}_2(\mathbb{Z})$ equivalence classes) can be composed. This led him to the phenomenon of the ideal class group before ideals were invented by Kummer and Dedekind. Besides in the ideal class group for more general number fields, Gauss's composition law has found an extension in Bhargava's higher composition laws. These are based on the representation theory of arithmetic groups ($\mathrm{SL}_2(\mathbb{Z})$ and its generalizations), in which regard they are natural structures in themselves. They have striking applications to old problems regarding mean asymptotics of Selmer ranks of elliptic curves, the $3$-parts of class groups of quadratic fields, etc.

*The Epstein zeta function takes the shape $\zeta_Q(s) := \sum_{\mathbf{n} \neq \mathbf{0}} Q(\mathbf{n})^{-s}$, for a given signature $(d,0)$ quadratic form $Q$. It has all the right analytical properties (meromorphic with simple pole at $s = 1$ and a functional equation relating $s \leftrightarrow 1-s$), allowing to decompose the zeta function of an imaginary quadratic field over a set of representatives $Q$ for the class group. This has consequences for the arithmetic of these fields, beautifully developed in Siegel's Lectures on Advanced Analytic Number Theory (Tata Institute lecture series, 1961). 

*For $d = 2$, $\zeta_Q(s)$ is in effect an Eisenstein series ($|mz+n|^2$ being a binary quadratic form in $m,n$), which is a natural structure all over mathematics, being a continuum of modular forms in the spectral resolution of the hyperbolic Laplacian. Siegel apparently had much interest in the conceptual role played in number theory by the higher rank quadratic forms and their Epstein zeta function. Much of his work was put on representation theoretic footing in Weil's 1964 paper Sur certains groupes d'operateurs unitaires. Michael Berg's book, The Fourier-Analytic Proof of Quadratic Reciprocity, is a terrific introduction to these ideas.
A: Perhaps too trivial an example; the fixed points of a (non-intentity) Mobius transform.
