The Bullet-Macdonald identity (c.f. On the Adem relations)is the following: $$P(s^2+st)P(t^2)=P(t^2+st)P(s^2).$$ where we have $P(s)=\Sigma _s s^iSq^i$. This is known to be equivalent to the Adem relations.
Now, this equality was proved with trivial action of Steenrod squares on the formal indeterminates in the paper (see also Larry Smith, An algebraic introduction to the Steenrod algebra, arXiv:0903.4997).
However, in the section 4 of the same paper, there is an alternative proof, where a different action was used in the course of the proof, but the conclusion is still about with the trivial action. There we have $Sq^1(t)=t^2$.
Another generating series identity equivalent to the Adem relations is the Bisson-Joyal identity, that is, we have $$Q(s)Q(t)=Q(t)Q(s)$$ with $Q(s)=\Sigma _s s^iSq_i$ where $Sq _i(x)=Sq^{n-i}(x)$ for $x\in H^n(X)$, and this time the Steenrod square acts non-trivially on the indeterminates by $$Q(s)(t)=t(s+t).$$
Since both Bullett-Macdonald and Bisson-Joyal identities are equivalent to Adem relations, they are equivalent to each other.
My questions are
Is there a direct proof of the equivalence between the two identities?
If there is any such proof, how does one go between the different actions of Steenrod squares?